• forewing vein R5 • stalked or fused with R3+4
• forewing without oval sclerotization costad of base of vein A1+2
• bullae tympani closed cephalad
• tympanum and conjunctivum in the same plane
• processus tympani absent
• praecinctorium absent
• accessory tympana absent
• male genitalia with uncus arms, (paired processes arising laterally from base of uncus)
• segment A8 of larvae almost always with sclerotized ring around base of SD1